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Ecological science has been shaped largely by the biological sciences. Environmental science, on the other hand, has been shaped largely by the physical sciences and engineering. With the beginning of interdisciplinary efforts between the two fields, some of the fundamental differences between them are generating conflicts caused more by misunderstanding of basic concepts than by any difference in social purposes or methods. Those differences are most vivid in that part of ecology called ecosystem science, for it is there that it is obvious that both the biota and the physical environment interact such that not only does the environment shape the biota but the biota transforms the environment. The accumulated body of empirical evidence concerning natural, disturbed, and managed ecosystems identifies key features of ecosystem structure and function (Holling et aI., 1995) that probably are not included in many engineers' image of ecology: